The paper was the outgrowth of observations mad on the Western plains. It is published in full in the American Naturalist for 1888. While recognizing the validity of protective, ornamental and typical coloration, as defined by Darwin, Wallace, Belt and others, the writer claimed that a very considerable amount of coloration in animals remained unexplained.
Proceedings of the Iowa Academy of Sciences
©1887 Iowa Academy of Science, Inc.
Todd, J. E.
"Directive Coloration in Animals,"
Proceedings of the Iowa Academy of Science, 1(Pt. 1), 14-15.
Available at: https://scholarworks.uni.edu/pias/vol1/iss1/11